Mitosis

Two major changes must occur

1) the condensation of the chromosomes

2) assembly of microtubules to form the spindle

Chromosomes consist of chromatids which become replicated during the S phase of the cell cycle and are joined together by a constricted area common to both chromatids. This region of attachment is known as the centromere. Disk-shaped protein-containing structures called kinetochores are sites to which the microtubule spindle fibers attach.

Spindle formation includes polar fibers and kinetochore fibers

Four stages of mitosis - nuclear division

Prophase -

  • centriole pairs begin to move to opposite poles of the cell.
  • nuclear envelope begins to disappear
  • chromosomes appear each with two chromatids.
  • spindle apparatus begins to assemble

Metaphase - chromosomes align at the equatorial plate.

Anaphase - individual chromatids are separated. The two sets of newly separated chromosomes are pulled by spindle fibers toward opposite poles of the cell.

Telophase - the nulcear envelope reappears. Two new nuclei are present. Chromosomes again begin to decondense becoming diffuse once more.

The spindle apparatus disappears.

Plasma membrane begins to pinch (cytokinesis).

 

Cytokinesis - cytoplasmic division - usually accompanies nuclear division- Mechanisms involves actin filaments acting as "purse strings" being pulled together.